Biology · Behavioral Ecology and Evolutionary Biology

In a monogamous haplodiploid social insect colony, sterile workers assist the queen in producing additional sisters instead of reproducing themselves. According to Hamilton's rule, altruistic behaviour is favoured when the genetic benefit to relatives (weighted by coefficient of relatedness, r) exceeds the cost, 'c' to the actor. If a worker helps the queen raise three daughters instead of producing two of her own, then based on Hamilton's rule, will helping behaviour evolve? [assume genetic unit as the number of offsprings weighted by r as a measure of inclusive fitness]